Holley, Robert William (1922–1993)
Robert Holley was a US biochemist who shared, with Har Khorana and Marshall Nirenberg, the 1968 Nobel
Prize in Physiology or Medicine for his work on the way genes determine the function of cells. Holley described
the first full sequence of sub-units in nucleic
acid, the genetic material of a cell. This was an important step toward
the understanding of gene action.